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A Monograph on the Sub-class Cirripedia, Volume 1, a non-fiction book by Charles Darwin |
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4. Oxynaspis |
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_ OXYNASPIS.[33] Gen. Nov. Pl. III. Valvæ 5, approximatæ: scutorum umbones in medio marginis occludentis positi: carina rectangulè flexa, sursùm inter terga extensa, termino basali simpliciter concavo. Valves 5, approximate; scuta with their umbones in the middle of the occludent margin; carina rectangularly bent, extending up between the terga, with the basal end simply concave. [33] From [Greek: oxunô], to sharpen, and [Greek: aspis], a shield or scutum.Mandibles with four teeth; maxillæ notched, with the lower part of edge nearly straight, prominent; anterior ramus of the second cirrus thicker than the posterior ramus; caudal appendages, uniarticulate, spinose. Attached to horny corallines.I have most unwillingly instituted this genus; but it will be seen by the following description, that the one known species could not have been introduced into Lepas or Pæcilasma, without destroying these genera, although it has a close general resemblance with both. As far as the valves are concerned, it is more nearly related to Lepas than to Pæcilasma; but taking the entire animal, its relation is much closer to the latter genus than to Lepas: it differs from both these genera in the manner of growth of the scuta, which is both upwards and downwards, the primordial valve being situated in nearly the middle of the occludent margin. In this respect, and in the shape of the carina and terga, there is an almost absolute identity with Scalpellum; I may, however, remark that in Scalpellum, the scuta first grow downwards, and afterwards in most of the species upwards, whereas here from the beginning, the growth is both upwards and downwards. In the mouth and cirri, there is rather more resemblance to Scalpellum than to Pæcilasma and Lepas: in habits, also, this genus agrees with Scalpellum, and if it had possessed a lower whorl of valves, it would have quite naturally entered that genus. It is unfortunate, that so insignificant and poorly characterised a form should require a generic appellation. In natural position, it appears to lead from Scalpellum through Pæcilasma to Lepas. 1. OXYNASPIS CELATA. Pl. III, fig. 1. Madeira; attached in numbers to an Antipathes; Rev. R. T. Lowe. Mus., Hancock.General Appearance.--The capitulum is rather thin, and broad in proportion to its length; it seems always entirely covered by the horny muricated bark of the Antipathes, and hence externally is coloured rich brown and covered with little horny spines. The membrane over the valves is very thin, and is with difficulty separated from the Antipathes; it has, I believe, no spines of its own. The corium lining the peduncle is a fine purple. All the individuals are attached to the coralline, with their capitulums upwards in the direction of the branches, and in this respect fig. 1. is erroneous. The valves, when cleared of the bark, are white, or are strongly tinged with pinkish-orange. The upper parts of the scuta and terga are plainly furrowed in lines radiating from their umbones; hence their margins are serrated with blunt teeth; their surfaces, moreover, are sparingly studded with small calcareous points. Scuta (fig. 1, a), sub-triangular, with the lower part rounded and protuberant, the upper produced and pointed. The umbo is situated in the middle of the occludent margin, instead of at the rostral angle, as in the foregoing genera. The occludent margin is straight, and is bordered by a narrow step or ledge, formed of transverse growth-ridges, and therefore has its edge serrated: the rostral angle is often slightly produced into a small projection. The basal margin is short, and forms an angle above a rectangle with the occludent margin: the tergal margin is straight; the carinal margin is rounded, protuberant, and of unusual length compared to the basal margin. The surface of the valve is convex near the umbo; and beneath there is a large deep hollow for the adductor muscle. Terga (fig. 1, b) large, flat, triangular, as long as the scuta or the carina, all three valves being nearly equal in length; occludent margin straight, or slightly arched, basal angle broad, not very sharp. Carina short (fig. 1, c, drawn rather too long), deeply concave, rectangularly bent, with the lower part not quite as long as the upper, and a little wider: the basal margin is truncated, rounded, and slightly sinuous. The umbo is situated at the angle, and therefore nearly central. The umbo of the terga, I may add, is in the same place, as in Lepas. The Peduncle is very short and narrow, and is, I believe, without spines; it is enveloped by the bark of the Antipathes. The capitulum in the largest specimens was .2 of an inch in length. Filamentary Appendages, apparently none. Mouth, with the orifice rather inclined abdominally. Labrum, with the upper part extremely protuberant, forming a projecting horn; no teeth on the crest. Palpi rather small, with only a few bristles at the end. Mandibles, with four teeth and the inferior angle pointed: first tooth as far from the second, as is the latter from the inferior angle; in one specimen, on one side, there were five teeth. Maxillæ, with three great spines at the upper angle, beneath which a deep notch, and with the inferior part much upraised; this lower part rather rounded at both corners, with the upper spines longer than the lower. Outer Maxillæ, with the bristles continuous in front; externally, slightly protuberant, with a tuft of bristles longer than those on the front side. Olfactory orifices apparently not protuberant; but all the specimens were in a bad state. Cirri.--Prosoma very little developed. First cirrus very far removed from the second. The three posterior cirri are straight and long; the segments are elongated and bear four or five pairs of very long spines, with a single minute intermediate spine between each pair; dorsal tufts, with long spines. First cirrus, rami unequal by two or three segments, and thickly covered with spines; the first cirrus is short compared to the second, owing to the length of the pedicel of the latter, though the longer ramus of the first, nearly equals the shorter ramus of the second pair. Second cirrus, with its anterior ramus shorter by two or three segments than the posterior ramus, and thicker than it, with the segments covered like brushes with bristles; posterior ramus, and both rami of the third cirrus, a little more thickly clothed with bristles than are the three posterior cirri. Caudal Appendages, minute, broadly oval, with six or seven long bristles on their summits. _ Read next: 5. Conchoderma Read previous: 3. 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